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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1021	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Nesonycteris woodfordi	Melonycteris woodfordi	Melonycteris woodfordi	Melonycteris woodfordi	Melonycteris woodfordi	Melonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi	Melonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi	Nesonycteris woodfordi		[MSW2] Subgenus Nesonycteris.; [MSW3] Subgenus Nesonycteris. Includes aurantius; see Flannery (1993b). Also see Flannery (1995b) and Bonaccorso (1998).; [HMW] Nesonycteris woodfordi Thomas, 1887 , “Shortland and Fauro Islands, western part of Solomon group [= Islands].” Nesonycteris woodfordi and N. fardoulisi might not be reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between them needs to be addressed with robust molecular and morphometric data; thorough revision of Nesonycteris is needed. Two subspecies recognized.; [batnames2022] Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius ; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).; [MDD2022] moved from Melonycteris to Nesonycteris.; [IUCN] The species Melonycteris aurantius was synonymized with this species.; [batnames2023] Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius ; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).; [MDD2023] includes the name schouteni, which was initially considered a subspecies of N. fardoulisi; moved from Melonycteris to Nesonycteris.; [MDD2025_2.0] includes the name schouteni, which was initially considered a subspecies of N. fardoulisi; moved from Melonycteris to Nesonycteris.; [batnames2025_1.7] Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).; [MDD2025_2.2] includes the name schouteni, which was initially considered a subspecies of N. fardoulisi; moved from Melonycteris to Nesonycteris.								woodfordi, aurantius		woodfordi, aurantius		woodfordi, aurantius		woodfordi, aurantius	The species Melonycteris aurantius was synonymized with this species.	woodfordi, aurantius		woodfordi, aurantius , schouteni	woodfordi, aurantia, schouteni	aurantius, woodfordi 		woodfordi O. Thomas, 1887|aurantia (C. J. Phillips, 1966)|schouteni (Flannery, 1993)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Woodford's fruit bat	Solomons	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Melonycteris woodfordi	Solomon Isis., Alu Isl. (near Shortland island).	Thomas	1887	Ann. Mag. Nat. Hist., 19:147.	Distribution: Known from a number of islands from Bougainville to Guadalcanal in the Solomons.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Woodford's fruit bat	Solomons	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1887	Ann. Mag. Nat. Hist., ser. 5, 19:147.	Subgenus Nesonycteris.	Solomon Isis.	Solomon Isis, Alu Isl (near Shortland Isl).		THOMAS	1887	Postorbital re gion of skull relatively narrow (7.2-7.5 mm). Size medium (forearm length, 52-58 mm). Pelage dark brown, little contrast between dorsum and ventrum.	Distribution: Known from a number of islands from Bougainville to Guadalcanal in the Solomons.	No subspecies.		39	species	M. woodfordi	THOMAS	1887	Nesonycteris	subgenus	Melonycteris woodfordi				Postorbital re gion of skull relatively narrow (7.2-7.5 mm). Size medium (forearm length, 52-58 mm). Pelage dark brown, little contrast between dorsum and ventrum.	No subspecies.		3. M. woodfordi (THOMAS 1887).	3	_N. w. aurantia_ (Phillips, 1966); _N. w. schouteni_ (Flannery, 1993); _N. w. woodfordi_ Thomas, 1887			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Melonycteris woodfordi	Melonycteris	Nesonycteris	woodfordi	Thomas	y	1887		Ann. Mag. Nat. Hist., ser. 5	19		147		Woodford's Fruit Bat	Solomon Isls, Western Province, Alu Isl (near Shortland Isl)	Bougainville and Buka Isls (Papua New Guinea), N and W Solomon Isls.	IUCN/SSC Action Plan (1992) – Not Threatened as M. woodfordi, No Data: Limited Distribution as M. aurantius. IUCN 2003 – Lower Risk (lc) as M. woodfordi; Vulnerable as M. aurantius.	aurantius Phillips, 1966.	Subgenus Nesonycteris. Includes aurantius; see Flannery (1993b). Also see Flannery (1995b) and Bonaccorso (1998).	03AD87FAFF8CF6628CB331E9F89EF210	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	130	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF8CF6628CB331E9F89EF210.xml	Nesonycteris woodfordi	Pteropodidae	Nesonycteris	woodfordi	Thomas	1887	Mélonyctére de Woodford @fr | Woodford-Blitenflughund @de | Melonicterio de Woodford @es | \Voodford's Fruit Bat @en	Nesonycteris woodfordi Thomas, 1887 , “Shortland and Fauro Islands, western part of Solomon group [= Islands].” Nesonycteris woodfordi and N. fardoulisi might not be reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between them needs to be addressed with robust molecular and morphometric data; thorough revision of Nesonycteris is needed. Two subspecies recognized.	N.w.woodfordiThomas,1887—SolomonIs(Buka,Bougainville,Shortlands,Fauro,Choiseul,andSantaIsabel). N. w. aurantius C. J. Phillips, 1966 — Solomon Is (Nggela).	Head—-body 80-106 mm (tailless), ear 10-16 mm , hindfoot 14- 4— 23 mm , forearm 48-59 mm ; weight 25-50 g . Subspecies aurantiusis smaller than nominate woodfordi and has distinct cranial measurements. Head is elongated, and rostrum is long, with tubular nostrils. Ears are relatively short and bluntly pointed; eyes are large, with dark brown irises. Dorsal pelage is orange to cinnamon, with pale russet head and nape; ventral pelage is pale grayish brown. Wings are black, with white on and surrounding digits; second digit is missing characteristic claw of most pteropodids. Uropatagium is extremely underdeveloped and barely present along inner leg down to ankle. Claws are brown, with white tips. Skull has long and narrow rostrum, with less well-developed (or completely absent in subspecies aurantius) sagittal crest in males unlike in Fardoulis’s Blossom Bat (N. fardoulisi ). 1, is missing, P is larger than M,, and molars and premolars are largely reduced and flattened.	Primary tropical forests but commonly secondary forests, rural gardens, and coconut plantations from sea level up to elevations of ¢. 1100 m .	Woodford’s Blossom Batis nectarivorous, probably feeding mostly on nectar, pollen, and occasionally soft fruit.	Lactating and pregnant Woodford’s Blossom Bats have been captured in February-March, May, July, September, and November, suggesting year-round breeding. Litters have one young. Apparently, females never carry dependent young, leaving them in roosts.	Woodford’s Blossom Bats are nocturnal, and day roosts are in foliage.	Woodford’s Blossom Bats seem to roost alone or in small groups and have relatively small home ranges.	Classified as Least Concern on The IUCN Red List (as Melonycteris woodfordi ). Woodford’s Blossom Bat is poorly known but seems to tolerate human disturbance. Threats are not currently known, but commercial logging might be a threat.	Bonaccorso (1998) | Flannery (1993, 1995a) | Giannini & Simmons (2007a) | Lavery (2017b) | Phillips (1966) | Pulvers & Colgan (2007)		126. Woodford’s Blossom Bat Nesonycteris woodfordi French: Mélonyctére de Woodford / German: Woodford-Blitenflughund / Spanish: Melonicterio de Woodford Other common names: \Voodford's Fruit Bat Taxonomy. Nesonycteris woodfordi Thomas, 1887 , “Shortland and Fauro Islands, western part of Solomon group [= Islands].” Nesonycteris woodfordi and N. fardoulisi might not be reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between them needs to be addressed with robust molecular and morphometric data; thorough revision of Nesonycteris is needed. Two subspecies recognized. Subspecies and Distribution. N.w.woodfordiThomas,1887—SolomonIs(Buka,Bougainville,Shortlands,Fauro,Choiseul,andSantaIsabel). N. w. aurantius C. J. Phillips, 1966 — Solomon Is (Nggela). Descriptive notes. Head—-body 80-106 mm (tailless), ear 10-16 mm , hindfoot 14- 4— 23 mm , forearm 48-59 mm ; weight 25-50 g . Subspecies aurantiusis smaller than nominate woodfordi and has distinct cranial measurements. Head is elongated, and rostrum is long, with tubular nostrils. Ears are relatively short and bluntly pointed; eyes are large, with dark brown irises. Dorsal pelage is orange to cinnamon, with pale russet head and nape; ventral pelage is pale grayish brown. Wings are black, with white on and surrounding digits; second digit is missing characteristic claw of most pteropodids. Uropatagium is extremely underdeveloped and barely present along inner leg down to ankle. Claws are brown, with white tips. Skull has long and narrow rostrum, with less well-developed (or completely absent in subspecies aurantius) sagittal crest in males unlike in Fardoulis’s Blossom Bat (N. fardoulisi ). 1, is missing, P is larger than M,, and molars and premolars are largely reduced and flattened. Habitat. Primary tropical forests but commonly secondary forests, rural gardens, and coconut plantations from sea level up to elevations of ¢. 1100 m . Food and Feeding. Woodford’s Blossom Batis nectarivorous, probably feeding mostly on nectar, pollen, and occasionally soft fruit. Breeding. Lactating and pregnant Woodford’s Blossom Bats have been captured in February-March, May, July, September, and November, suggesting year-round breeding. Litters have one young. Apparently, females never carry dependent young, leaving them in roosts. Activity patterns. Woodford’s Blossom Bats are nocturnal, and day roosts are in foliage. Movements, Home range and Social organization. Woodford’s Blossom Bats seem to roost alone or in small groups and have relatively small home ranges. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Melonycteris woodfordi ). Woodford’s Blossom Bat is poorly known but seems to tolerate human disturbance. Threats are not currently known, but commercial logging might be a threat. Bibliography. Bonaccorso (1998), Flannery (1993, 1995a), Giannini & Simmons (2007a), Lavery (2017b), Phillips (1966), Pulvers & Colgan (2007).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Nesonycteris woodfordi	Nesonycteris		woodfordi	Thomas	1887	0	Proc. Zool. Soc. London	1892:24:00	Woodford's Fruit Bat	<b> aurantius </b> Phillips, 1966.	Solomon Isls, Western Province, Alu Isl (near Shortland Isl)	Bougainville and Buka Isls (Papua New Guinea), N and W Solomon Isls	Not listed.	Least Concern as Melonycteris woodfordi 	Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius ; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nesonycteris woodfordi	23	Woodford's Blossom Bat	Woodford's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	MELONYCTERINI	Nesonycteris	NA	woodfordi	O. Thomas	1887	0	Nesonycteris_woodfordi	Thomas, O. (1887). Diagnoses of two new fruit-eating bats from the Solomon Islands. Annals and Magazine of Natural History, ser. 5, 19, 147.	https://www.biodiversitylibrary.org/item/55145#page/161/mode/1up	BM 1887.1.18.9		"Shortland and Fauro Islands, western part of Solomon group [= Islands]."			woodfordi (O. Thomas, 1887)|aurantius C. J. Phillips, 1966	moved from Melonycteris to Nesonycteris.	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.	Papua New Guinea|Solomon Islands	Oceania	Australasia/Oceania	LC	0	0	0	Nesonycteris_woodfordi	0	sciname match	Melonycteris_woodfordi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13140	Melonycteris woodfordi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Melonycteris	woodfordi	Thomas, 1887	The species Melonycteris aurantius was synonymized with this species.	20000000	Melonycteris woodfordi	Least Concern		2017	2016-06-13 00:00:00 UTC	3.1	English	Listed as Least Concern because, its extent of occurrence and area of occupancy are large, and it is commonly recorded in a wide range of habitats including disturbed secondary forests. Although population numbers are thought to be large, there are few data available toconfirm this or identify trends.	It is known from primary tropical forest, but is also commonly found in disturbed habitats including secondary forest, rural gardens, and coconut plantations (Bonaccorso 1998, Flannery 1995). It roosts in foliage in small groups or individually and likely forages over a relatively small home range (S. Hamilton pers. comm.).	Threats are poorly known, habitat loss due to commercial logging may adversely affect the species. Because this species does not require caves and does not roost in large colonies, it is immune to many of the threats faced by cave roosting bats of the region (S. Hamilton pers. comm.). However, a sister species M. melanops has been shown to forage over relatively short home ranges (Bonaccorso and Winkelmann 2005). If M. woodfordi forages over a similarly small home range this may render it vulnerable to local environmental disturbances such as commercial logging.	It is a species that is commonly captured in mist nets on Choiseul, Bougainville and Santa Isabel. Few data are available to determine population sizes or trends. Detailed studies of the sister species M. fardoulisi have shown dramatic declinesamong the New Georgia group of islands. It is assumed M. woodfordi fills a similar ecological niche to this species and as such may be similarly vulnerable to environmental perturbations. Melonycteris woodfordi is often recorded in degraded habitats, but long-term population data mayshow it qualifies for listing in a threatened category.	Unknown	This species is present in Solomon Island chain (Papua New Guinea and Solomon Islands). The subspecies Melonycteris woodfordi woodfordi is found on the islands of Bougainville and Buka of Papua New Guinea, and the islands of Oblari, Alu, Choiseul, Fauro, Mono, and Santa Isabel in the Solomon Islands. The subspecies M. w. aurantius is found on Nggela and Florida Islands in the Solomon Islands (Bonaccorso 1998, Flannery 1995). The species occurs from sea level to 1,100 m asl.		Terrestrial	The population numbers and trends of this species should be assessed.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Nesonycteris		woodfordi	Thomas	1887	0	Proc. Zool. Soc. London	1892:24:00	Woodford's Fruit Bat	<b> aurantius </b> Phillips, 1966.	Solomon Isls, Western Province, Alu Isl (near Shortland Isl)	Bougainville and Buka Isls (Papua New Guinea), N and W Solomon Isls	Not listed.	Least Concern as Melonycteris woodfordi 	Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius ; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).	Nesonycteris woodfordi	1004433	23	Woodford's Blossom Bat	Woodford's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	MELONYCTERINI	Nesonycteris	NA	woodfordi	O. Thomas	1887	0	Nesonycteris_woodfordi	Thomas, O. (1887). Diagnoses of two new fruit-eating bats from the Solomon Islands. Annals and Magazine of Natural History, ser. 5, 19, 147.	https://www.biodiversitylibrary.org/item/55145#page/161/mode/1up	BM 1887.1.18.9		"Shortland and Fauro Islands, western part of Solomon group [= Islands]."			woodfordi O. Thomas, 1887|aurantius (C. J. Phillips, 1966)|schouteni (Flannery, 1993)	includes the name schouteni, which was initially considered a subspecies of N. fardoulisi; moved from Melonycteris to Nesonycteris.	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.|Lavery, T. H., DeCicco, L. H., Olson, K. V., Holland, P. S., & Moyle, R. G. (2023). Phylogeography of Solomon Islands blossom bats reflects oceanic divides and Pleistocene connections. Journal of Biogeography, 00, 1-12.				Papua New Guinea|Solomon Islands	Oceania	Australasia/Oceania	LC	0	0	0	Nesonycteris_woodfordi	0	sciname match	Melonycteris_woodfordi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nesonycteris_woodfordi	1004433	23	Woodford's Blossom Bat	Woodford's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Melonycterini	Nesonycteris	NA	woodfordi	O. Thomas	0	Nesonycteris woodfordi	Thomas, O. 1887-02-01. Diagnoses of two new fruit-eating bats from the Solomon Islands. Annals and Magazine of Natural History (5)19(110):147.	https://www.biodiversitylibrary.org/page/16334742	BMNH:Mamm:1887.1.18.9	holotype	https://data.nhm.ac.uk/object/c00179b7-6e79-41bb-aa65-10870b9acef9	"Shortland and Fauro Islands, western part of Solomon group [= Islands]."			includes the name schouteni, which was initially considered a subspecies of N. fardoulisi; moved from Melonycteris to Nesonycteris.	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.|Lavery, T. H., DeCicco, L. H., Olson, K. V., Holland, P. S., & Moyle, R. G. (2023). Phylogeography of Solomon Islands blossom bats reflects oceanic divides and Pleistocene connections. Journal of Biogeography, 00, 1-12.				Papua New Guinea|Solomon Islands	Oceania (Continent)	Australasia	LC	0	0	0	Nesonycteris_woodfordi	0	sciname match	Melonycteris_woodfordi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Nesonycteris		woodfordi	Thomas	1887	0	Proc. Zool. Soc. London	1892:24:00	Woodford's Fruit Bat	aurantius Phillips, 1966.	Solomon Isls, Western Province, Alu Isl (near Shortland Isl)	Bougainville and Buka Isls (Papua New Guinea), N and W Solomon Isls, including Guadalcanal	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13140/21977332/' target='_blank'>Least Concern</a>	Transferred to Nesonycteris by Almeida et al. (2020). Includes aurantius; see Flannery (1993). Also see Flannery (1995) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Melonycteris woodfordi; Nesonycteris woodfordi; Nesonycteris woodfordi; Nesonycteris woodfordi; Melonycteris woodfordi; Nesonycteris woodfordi; woodfordi; aurantius; woodfordi; aurantius; aurantius; woodfordi; aurantius; Mélonyctére de Woodford; Woodford-Blitenflughund; Melonicterio de Woodford; \Voodford's Fruit Bat; Woodford's Blossom Bat; Woodford's Fruit Bat; Woodford's Fruit Bat; Woodford's Fruit Bat; N. woodfordi