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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L103	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Barbastella pacifica	Barbastella pacifica	Barbastella pacifica	N/A	Barbastella pacifica	Barbastella pacifica	Barbastella pacifica	Barbastella pacifica	Barbastella pacifica		[HMW] Barbastella pacifica Kruskop, Kawai & Tiunov, 2019 , “Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°.” See B. darjelingensis . Monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														pacifica				pacifica	pacifica			pacifica Kruskop, Kawai, & M. P. Tiunov, 2019 [preoccupied]						N/A																																								NA																											4C3D87E8FF936A2CFF499EE51401BFA9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	862	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF936A2CFF499EE51401BFA9.xml	Barbastella pacifica	Vespertilionidae	Barbastella	pacifica	Kruskop, Kawai & Tiunov	2019	Barbastelle du Japon @fr | Japan-Mopsfledermaus @de | Barbastelade Japon @es	Barbastella pacifica Kruskop, Kawai & Tiunov, 2019 , “Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°.” See B. darjelingensis . Monotypic.	Kunashir I, Hokkaido , and scattered on Honshu and Shikoku, Japan .	Head-body 4761- 6 mm , tail 41-52 mm , ear 12-5—- 19 mm , hindfoot 6-6-10- 2 mm , forearm 39 43 mm . The Japanese Barbastelle is a large species of barbastelle that can be distinguished by its darker pelage (especially ventrally), braincase, mandible, and baculum. Pelage is long, thick, and soft with somewhat wavy underfur hairs; dorsally deep brown shadowed by light brown (less glossy than in the Caspian Barbastelle, B. caspica, and the Arabian Barbastelle, B. leucomelas ); ventrally similar to dorsum. Immatures are darker overall. Face is short with large inflations on either side of muzzle, as in other barbastelles. Naked portions of wings, face, and ears dark brown. Ears broadly triangular, thick and rounded at tip, with wrinkled upper mid-anterior margin, wrinkles extending into inner ear; ears connected at thick frontal point, are relatively long, and lack an additional lobe on outer margin (present in the Western Barbastelle, B. barbastellus , and the Beijing Barbastelle, B. beijingensis ); tragus long and more or less triangular, with slightly concave posterior margin in upper one-third and convex posterior margin at widest part. Wing attaches to base offirst toe; uropatagium stretches from ankle to tail tip; calcar reaches less than halfway to long tail. Baculum has proportionally wide shaft with widened and very blunt distal tip (wider and more massive than in the Eastern Barbastelle, B. darjelingensis ). Skull has moderately high, rounded braincase; frontal portion slightly higher than temporal portion but less so than in the Western Barbastelle, the Arabian Barbastelle, or the Beijing Barbastelle; a shallow depression can often be seen on the level of coronal suture; sagittal crest not developed; lambdoidal crests weak; angular process of mandible shorter and blunter than in the Beijing Barbastelle; facial profile moderately sloped; dental features are generally similar to other barbastelles. Chromosomal complement apparently has 2n = 32 and FN = 50.	Probably inhabits temperate mixed and broadleaf forests.	No information.	A female with a single embryo has been observed.	Nocturnal, foraging through the night and roosting by day. Has been reported roosting in artificial caves, tunnels, and rock crevices. Hibernates through winter.	No information.	Not assessed on The IUCN Red List. The species has a fairly restricted range across Japan , and seems to be relatively rare. Virtually nothing is known of its ecology and threats; further research is certainly needed.	Benda & Mlikovsky (2008) | Kruskop (2015) | Kruskop et al. (2019) | Ohdachi et al. (2015)	https://zenodo.org/record/6398281/files/figure.png	223. Japanese Barbastelle Barbastella pacifica French: Barbastelle du Japon / German: Japan-Mopsfledermaus / Spanish: Barbastela de Japon Taxonomy. Barbastella pacifica Kruskop, Kawai & Tiunov, 2019 , “Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°.” See B. darjelingensis . Monotypic. Distribution. Kunashir I, Hokkaido , and scattered on Honshu and Shikoku, Japan . Descriptive notes. Head-body 4761- 6 mm , tail 41-52 mm , ear 12-5—- 19 mm , hindfoot 6-6-10- 2 mm , forearm 39 43 mm . The Japanese Barbastelle is a large species of barbastelle that can be distinguished by its darker pelage (especially ventrally), braincase, mandible, and baculum. Pelage is long, thick, and soft with somewhat wavy underfur hairs; dorsally deep brown shadowed by light brown (less glossy than in the Caspian Barbastelle, B. caspica, and the Arabian Barbastelle, B. leucomelas ); ventrally similar to dorsum. Immatures are darker overall. Face is short with large inflations on either side of muzzle, as in other barbastelles. Naked portions of wings, face, and ears dark brown. Ears broadly triangular, thick and rounded at tip, with wrinkled upper mid-anterior margin, wrinkles extending into inner ear; ears connected at thick frontal point, are relatively long, and lack an additional lobe on outer margin (present in the Western Barbastelle, B. barbastellus , and the Beijing Barbastelle, B. beijingensis ); tragus long and more or less triangular, with slightly concave posterior margin in upper one-third and convex posterior margin at widest part. Wing attaches to base offirst toe; uropatagium stretches from ankle to tail tip; calcar reaches less than halfway to long tail. Baculum has proportionally wide shaft with widened and very blunt distal tip (wider and more massive than in the Eastern Barbastelle, B. darjelingensis ). Skull has moderately high, rounded braincase; frontal portion slightly higher than temporal portion but less so than in the Western Barbastelle, the Arabian Barbastelle, or the Beijing Barbastelle; a shallow depression can often be seen on the level of coronal suture; sagittal crest not developed; lambdoidal crests weak; angular process of mandible shorter and blunter than in the Beijing Barbastelle; facial profile moderately sloped; dental features are generally similar to other barbastelles. Chromosomal complement apparently has 2n = 32 and FN = 50. Habitat. Probably inhabits temperate mixed and broadleaf forests. Food and Feeding. No information. Breeding. A female with a single embryo has been observed. Activity patterns. Nocturnal, foraging through the night and roosting by day. Has been reported roosting in artificial caves, tunnels, and rock crevices. Hibernates through winter. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. The species has a fairly restricted range across Japan , and seems to be relatively rare. Virtually nothing is known of its ecology and threats; further research is certainly needed. Bibliography. Benda & Mlikovsky (2008), Kruskop (2015), Kruskop et al. (2019), Ohdachi et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Barbastella pacifica	Barbastella		pacifica	Kruskop, Kawai & Tiunov	2019	0	Zootaxa	4567(3): 470	Japanese Barbastelle	None.	Russia, Kunashir (Kunashiri Island), Filatova (Ruyabetsu) stream, ca. 1 km from the mouth	Russia (Kuril Isls.), Japan	Not listed - new species.	Not Evaluated		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Barbastella pacifica	23	Japanese Barbastelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PLECOTINI	Barbastella	NA	pacifica	Kruskop, Kawai, & Tiunov	2019	0	Barbastella_pacifica	Kruskop, S. V., Kawai, K., & Tiunov, M. P. (2019). Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa, 4567(3), 470.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4567.3.3			"Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°."	44.19	146.02	pacifica Kruskop, Kawai, & Tiunov, 2019	recently described	Kruskop, S. V., Kawai, K., & Tiunov, M. P. (2019). Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa, 4567(3), 461-476.	Japan	Asia	Palearctic	NA	0	0	0	Barbastella_pacifica	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Barbastella		pacifica	Kruskop, Kawai & Tiunov	2019	0	Zootaxa	4567(3): 470	Japanese Barbastelle	None.	Russia, Kunashir (Kunashiri Island), Filatova (Ruyabetsu) stream, ca. 1 km from the mouth	Russia (Kuril Isls.), Japan	Not listed - new species.	Not Evaluated		Barbastella pacifica	1005654	23	Japanese Barbastelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PLECOTINI	Barbastella	NA	pacifica	Kruskop, Kawai, & Tiunov	2019	0	Barbastella_pacifica	Kruskop, S. V., Kawai, K., & Tiunov, M. P. (2019). Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa, 4567(3), 470.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4567.3.3			"Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°."	44.19	146.023	pacifica Kruskop, Kawai, & Tiunov, 2019	recently described	Kruskop, S. V., Kawai, K., & Tiunov, M. P. (2019). Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa, 4567(3), 461-476.				Japan	Asia	Palearctic	NA	0	0	0	Barbastella_pacifica	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Barbastella_pacifica	1005654	23	Japanese Barbastelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Plecotini	Barbastella	NA	pacifica	Kruskop, Kawai, & M. P. Tiunov	0	Barbastella pacifica	Kruskop, S.V., Kawai, K. and Tiunov, M.P. 2019-03-18. Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa 4567(3):461-476.	https://doi.org/10.11646/zootaxa.4567.3.3	IBSS 168-07	holotype		"Kunashir (Kunashiri) Island, Filatova (Ruyabetsu) stream, ca. 1 km from the mouth (Selezneva & Tiunov 2007); N 44.190°, E 146.023°."	44.19	146.023	recently described	Kruskop, S. V., Kawai, K., & Tiunov, M. P. (2019). Taxonomic status of the barbastelles (Chiroptera: Vespertilionidae: Barbastella) from the Japanese archipelago and Kunashir Island. Zootaxa, 4567(3), 461-476.				Japan	Asia	Palearctic	NE	0	0	0	Barbastella_pacifica	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Barbastella		pacifica	Kruskop, Kawai & Tiunov	2019	0	Zootaxa	4567(3): 470	Japanese Barbastelle	None.	Russia, Kunashir (Kunashiri Island), Filatova (Ruyabetsu) stream, ca. 1 km from the mouth	Russia (Kuril Isls.), Japan	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Barbastella pacifica; Barbastella pacifica; Barbastella pacifica; Barbastella pacifica; pacifica; Barbastelle du Japon; Japan-Mopsfledermaus; Barbastelade Japon; Japanese Barbastelle; Japanese Barbastelle; B. pacifica