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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1018	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Melonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi	Melonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi	Nesonycteris fardoulisi		[MSW3] Subgenus Nesonycteris. See Flannery (1993b, 1995b).; [HMW] Melonycteris fardoulisi Flannery, 1993 , “secondary forest near Sesena Village ( 8°31’S 162°05’E ), altitude 100- 9200 m , Makira |= San Cristobal Island ], Solomon Islands .” Nesonycteris is traditionally considered a subgenus of Melonycteris ; however, the two are considered distinct genera here based on their high degree of morphological distinctions, including distinct dentition, pelage pattern, and presence of index claw, and their relatively high degree of genetic differentiation. Nesonycteris woodford and N. fardoulisi are not reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between these two species needs to be addressed with robust molecular and morphometric data. Four subspecies recognized.; [batnames2022] Transferred to Nesonycteris by Almeida et al. (2020). See Flannery (1993, 1995).; [MDD2022] moved from Melonycteris to Nesonycteris.; [batnames2023] Transferred to Nesonycteris by Almeida et al. (2020). See Flannery (1993, 1995).; [MDD2023] previously included N. maccoyi and N. mengermani; moved from Melonycteris to Nesonycteris; [MDD2025_2.0] previously included N. maccoyi and N. mengermani; moved from Melonycteris to Nesonycteris; [batnames2025_1.7] Previously considered a subgenus of Melonycteris (see e.g., Phillips, 1968; Flannery 1993; Simmons, 2005). Raised to full generic status by Almeida et al. (2020). Melonycteris, including Nesonycteris, was revised by Flannery (1993). Does not include maccoyi or mengermani; see Pulvers and Colgan (2006) and Lavery et al. (2023). See also Flannery (1995). Populations on Guadalcanal previously attributed to fardoulisi appear to belong to woodfordi; see Lavery et al. (2023). For phylogeography see Pulvers and Colgan (2006) and Lavery et al. (2023).; [MDD2025_2.2] previously included N. maccoyi and N. mengermani; moved from Melonycteris to Nesonycteris								fardoulisi, maccoyi, mengermani, schouteni		fardoulisi, maccoyi, mengermani, schouteni		fardoulisi, maccoyi, mengermani, schouteni		fardoulisi, maccoyi, mengermani, schouteni		fardoulisi, schouteni		fardoulisi	fardoulisi	fardoulisi, schouteni		fardoulisi (Flannery, 1993)						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Melonycteris fardoulisi	Melonycteris	Nesonycteris	fardoulisi	Flannery		1993		Rec. Aust. Mus.	45		68		Fardoulis's Blossom Bat	Solomon Isls., Makira (= San Cristobal Isl.), Sesena.	S and E Solomon Isls.	IUCN 2003 – Vulnerable.	maccoyi Flannery, 1993; mengermani Flannery, 1993; schouteni Flannery, 1993.	Subgenus Nesonycteris. See Flannery (1993b, 1995b).	03AD87FAFF8CF66289AF3C88F8ADFA36	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	130	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF8CF66289AF3C88F8ADFA36.xml	Nesonycteris fardoulisi	Pteropodidae	Nesonycteris	fardoulisi		1993	Mélonyctére de Fardoulis @fr | Fardoulis-Blitenflughund @de | Melonicterio de Fardoulis @es	Melonycteris fardoulisi Flannery, 1993 , “secondary forest near Sesena Village ( 8°31’S 162°05’E ), altitude 100- 9200 m , Makira |= San Cristobal Island ], Solomon Islands .” Nesonycteris is traditionally considered a subgenus of Melonycteris ; however, the two are considered distinct genera here based on their high degree of morphological distinctions, including distinct dentition, pelage pattern, and presence of index claw, and their relatively high degree of genetic differentiation. Nesonycteris woodford and N. fardoulisi are not reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between these two species needs to be addressed with robust molecular and morphometric data. Four subspecies recognized.	N.f.fardoulisiFlannery,1993—SolomonIs(Makira). N.f.maccoyiFlannery,1993—SolomonIs(Malaita). N.f.mengermaniFlannery,1993—SolomonIs(VellaLavella,Kolombangara,NewGeor-gia,Vangunu,andRussell). N. f. schouteni Flannery, 1993 — Solomon Is ( Guadalcanal ).	Head-body 87-96 mm (tailless), ear 11- 9-14 mm , forearm 57- 9- 61 mm ; weight 39- 5-54 g . Males seem to be larger than females, with larger canines and more robust heads, and overall pelage of males is darker than females. Subspecies vary in cranial features, overall size, and amount of pink spotting over much of body, with fardoulisi and maccoyi being largest and mengermani being smallest. Head of Fardoulis’s Blossom Batis elongated, and rostrum is long, with tubular nostrils. Ears are relatively short and bluntly pointed; eyes are large, with dark brown irises. Pelage is generally light (females) to dark (males) brown to cinnamon-brown; ventral pelage is somewhat lighter brown drab. Skin on wings, ears (only in subspecies fardoulisi ), face, and feet have variable amounts of bright pink spotting (absent in subspecies maccoyi). Wings are black, with pink digits and variable amounts of bright pink spotting; second digit of wing is lacking characteristic claw of most pteropodids. Uropatagium is completely absent. Claws are brown with white tips. Skull has long and narrow rostrum, with well-developed sagittal crests in males (unlike in Woodford’s Blossom Bat, N. woodfordi ) and long infraorbital canal. Dental formula for all species of Nesonycteris is 12/1, Cl/1,P 53/3, M2/3:02) = 32. | ismissing; P'is larger than M,; and molars and premolars are largely reduced and flattened.	Primary montane tropical forests, lowland forests, rural gardens, and disturbed areas. Male and female Fardoulis’s Blossom Bats might forage separately and within distinct habitats because males seem to be found more often in disturbed habitats, but this has not been confirmed.	Fardoulis’s Blossom Batis nectarivorous, most likely eating nectar, pollen, and occasionally some soft fruits. It will feed on nectar of Heliconia solomonensis ( Heliconiaceae ), visiting plants in sequential fashion (trap-lining behavior); it is the sole pollinator ofthis self<incompatible plant. It lands on an inflorescence and clings to it using feet and thumb claws. Nectar is taken from open flowers, and pollen is deposited on chin and neck.	No information.	Fardoulis’s Blossom Bat is nocturnal and might roost in foliage.	Fardoulis’s Blossom Bat seems to be solitary.	Classified as Least Concern on The IUCN Red List (as Melonycteris fardoulisi ). Fardoulis’s Blossom Bat is variably common or rare throughout its distribution. It is found in various disturbed areas and seems to tolerate human disturbance. Habitat destruction from logging and agricultural expansion might be its biggest threats in the future.	Bonaccorso, Helgen & Hamilton (2008a) | Flannery (1993) | Giannini & Simmons (2007a) | Kress (1985) | Pulvers & Colgan (2007)		125. Fardoulis’s Blossom Bat Nesonycteris fardoulisi French: Mélonyctére de Fardoulis / German: Fardoulis-Blitenflughund / Spanish: Melonicterio de Fardoulis Taxonomy. Melonycteris fardoulisi Flannery, 1993 , “secondary forest near Sesena Village ( 8°31’S 162°05’E ), altitude 100- 9200 m , Makira |= San Cristobal Island ], Solomon Islands .” Nesonycteris is traditionally considered a subgenus of Melonycteris ; however, the two are considered distinct genera here based on their high degree of morphological distinctions, including distinct dentition, pelage pattern, and presence of index claw, and their relatively high degree of genetic differentiation. Nesonycteris woodford and N. fardoulisi are not reciprocally monophyletic, with N. f. schouteni being closest to N. w. woodfordi . Relationship between these two species needs to be addressed with robust molecular and morphometric data. Four subspecies recognized. Subspecies and Distribution. N.f.fardoulisiFlannery,1993—SolomonIs(Makira). N.f.maccoyiFlannery,1993—SolomonIs(Malaita). N.f.mengermaniFlannery,1993—SolomonIs(VellaLavella,Kolombangara,NewGeor-gia,Vangunu,andRussell). N. f. schouteni Flannery, 1993 — Solomon Is ( Guadalcanal ). Descriptive notes. Head-body 87-96 mm (tailless), ear 11- 9-14 mm , forearm 57- 9- 61 mm ; weight 39- 5-54 g . Males seem to be larger than females, with larger canines and more robust heads, and overall pelage of males is darker than females. Subspecies vary in cranial features, overall size, and amount of pink spotting over much of body, with fardoulisi and maccoyi being largest and mengermani being smallest. Head of Fardoulis’s Blossom Batis elongated, and rostrum is long, with tubular nostrils. Ears are relatively short and bluntly pointed; eyes are large, with dark brown irises. Pelage is generally light (females) to dark (males) brown to cinnamon-brown; ventral pelage is somewhat lighter brown drab. Skin on wings, ears (only in subspecies fardoulisi ), face, and feet have variable amounts of bright pink spotting (absent in subspecies maccoyi). Wings are black, with pink digits and variable amounts of bright pink spotting; second digit of wing is lacking characteristic claw of most pteropodids. Uropatagium is completely absent. Claws are brown with white tips. Skull has long and narrow rostrum, with well-developed sagittal crests in males (unlike in Woodford’s Blossom Bat, N. woodfordi ) and long infraorbital canal. Dental formula for all species of Nesonycteris is 12/1, Cl/1,P 53/3, M2/3:02) = 32. | ismissing; P'is larger than M,; and molars and premolars are largely reduced and flattened. Habitat. Primary montane tropical forests, lowland forests, rural gardens, and disturbed areas. Male and female Fardoulis’s Blossom Bats might forage separately and within distinct habitats because males seem to be found more often in disturbed habitats, but this has not been confirmed. Food and Feeding. Fardoulis’s Blossom Batis nectarivorous, most likely eating nectar, pollen, and occasionally some soft fruits. It will feed on nectar of Heliconia solomonensis ( Heliconiaceae ), visiting plants in sequential fashion (trap-lining behavior); it is the sole pollinator ofthis self<incompatible plant. It lands on an inflorescence and clings to it using feet and thumb claws. Nectar is taken from open flowers, and pollen is deposited on chin and neck. Breeding. No information. Activity patterns. Fardoulis’s Blossom Bat is nocturnal and might roost in foliage. Movements, Home range and Social organization. Fardoulis’s Blossom Bat seems to be solitary. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Melonycteris fardoulisi ). Fardoulis’s Blossom Bat is variably common or rare throughout its distribution. It is found in various disturbed areas and seems to tolerate human disturbance. Habitat destruction from logging and agricultural expansion might be its biggest threats in the future. Bibliography. Bonaccorso, Helgen & Hamilton (2008a), Flannery (1993), Giannini & Simmons (2007a), Kress (1985), Pulvers & Colgan (2007).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Nesonycteris fardoulisi	Nesonycteris		fardoulisi	Flannery	1993	1	Rec. Aust. Mus.	46:08:00	Fardoulis' Blossom Bat	<b> maccoyi </b>Flannery, 1993;<b> mengermani </b>Flannery, 1993;<b> schouteni </b>Flannery, 1993.	Solomon Isls., Makira (= San Cristobal Isl.), Sesena	S and E Solomon Isls	Not listed.	Near Threatened as Melonycteris fardoulisi 	Transferred to Nesonycteris by Almeida et al. (2020). See Flannery (1993, 1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Nesonycteris fardoulisi	23	Fardoulis's Blossom Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	MELONYCTERINI	Nesonycteris	NA	fardoulisi	Flannery	1993	1	Melonycteris_fardoulsi	Flannery, T. F. (1993). Revision of the genus Melonycteris (Pteropodidae: Mammalia). Records of the Australian Museum, 45(1), 68.	https://journals.australian.museum/flannery-1993-rec-aust-mus-451-5980/	AM M.18833		"secondary forest near Sesena Village (8°31'S 162°05'E), altitude 100–200 m, Makira [= San Cristobal Island], Solomon Islands."	-8.517	162.08	fardoulisi Flannery, 1993|maccoyi Flannery, 1993|mengermani Flannery, 1993|schouteni Flannery, 1993	moved from Melonycteris to Nesonycteris.	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.	Solomon Islands	Oceania	Australasia/Oceania	NT	0	0	0	Nesonycteris_fardoulisi	0	sciname match	Melonycteris_fardoulisi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13141	Melonycteris fardoulisi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Melonycteris	fardoulisi	Flannery, 1993		20000000	Melonycteris fardoulisi	Near Threatened	A2a	2020	2019-07-10 00:00:00 UTC	3.1	English	This bat is listed as Near Threatened (nearly meets criterion A2a) as its global population is estimated to have declined by 25-30% over the last 12.9 years (three generations; generation length = 4.3 years, Pacifici et al. 2013) as its habitat has been significantly degraded. In some locations (e.g. high elevation Malaita) it remains an extremely abundant species, being the most common bat captured in mist net surveys. However, surveys in Western Province, Solomon Islands show that overall capture rates in lowland forests of Kolombangara, New Georgia and Vangunu Islands may have declined by as much as 80% since 1992. Assuming a consistent rate of decline throughout this period, the overall capture rate in the Western Province may have declined by approximately 61% in the past 10 years. High molecular divergence and distinct morphology among island populations may warrant elevation of subspecies to full species and alter conservation status.	It is most often recorded in primary lowland forest but is also regularly encountered in other habitats (e.g., montane forest on Mount Makarakomburu, Guadalcanal; secondary forests; and village gardens). Males and females are sexually dimorphic and have distinctive dentition suggesting that foraging strategies may differ between the sexes, or strong competition between males for mates. Males have larger canines (Flannery 1993). Both sexes are aggressive if handled, and appear to be sedentary and territorial. It roosts in foliage (e.g cut nut Barringtonia edulis ) and is a common visitor to the flowers of bananas and wild ginger (Alpinia spp.) (T.H. Lavery pers. obs). In early November a juvenile animal that was unable to fly was discovered in a cut nut tree (Barringtonia edulis ) in village garden. It had presumably been deposited there whilst the mother was foraging (T.H. Lavery pers. obs).	Although this species appears able to persist in a variety of habitats, habitat degradation due to commercial logging and agricultural expansion probably poses a major threat to this species throughout across the majority of its range.	The species is widespread and remains a relatively commonly captured species in mist-net surveys (T.H. Lavery pers. obs.). However, recent comparisons of past and present mist-net capture rates in the Solomon Islands Western Province indicated substantial population declines between the years 1992 and 2015. Surveys were repeated at multiple sites across eight localities on the islands of Kolombangara (two localities), New Georgia (three localities) and Vangunu (three localities). Across these eight localities total capture rates have declined from an average of 62.62 individuals per 1,000 metre-squared mist net hours (m<sup>2</sup>net ;h<sup>-1</sup>) in 1992 to an average 13.45 per 1,000 m<sup>2</sup>net ;h<sup>-1</sup> in 2015. The total capture rate has declined from 4.02 individuals per 1,000 m<sup>2</sup>net ;h<sup>-1</sup> in 1992 to 0.82 individuals per 1,000 m<sup>2</sup>net ;h<sup>-1</sup> in 2015 (a decline of 80%). At one locality on New Georgia an increase in capture rates was recorded, and the species was not captured at one locality on Kolombangara in either survey period. At the remaining six localities, capture rates have declined by an average of almost 70%. This trend has been consistent regardless of land-use changes (i.e. the six localities are a mixture of primary forest, and forests that have been commercially logged before or after 1992). However, unlogged sites also lay in close proximity to commercial logging. Flannery (1995) reported that it was moderately common at some localities, but quite rare at others.	Decreasing	This species is endemic to the Solomon Islands. It has been recorded from Guadalcanal, Kolombangara, Malaita, New Georgia, the Russell Islands, San Cristobal, Vangunu, Gatokae, Ranongga, Tetepare and Vella Lavella (Flannery 1995, T. Lavery pers. obs.).		Terrestrial	Further studies are needed into the population status, distribution, and habitat requirements of this species.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Nesonycteris		fardoulisi	Flannery	1993	1	Rec. Aust. Mus.	46:08:00	Fardoulis' Blossom Bat	<b> maccoyi </b>Flannery, 1993;<b> mengermani </b>Flannery, 1993;<b> schouteni </b>Flannery, 1993.	Solomon Isls., Makira (= San Cristobal Isl.), Sesena	S and E Solomon Isls	Not listed.	Near Threatened as Melonycteris fardoulisi 	Transferred to Nesonycteris by Almeida et al. (2020). See Flannery (1993, 1995).	Nesonycteris fardoulisi	1004432	23	Fardoulis's Blossom Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	MELONYCTERINI	Nesonycteris	NA	fardoulisi	Flannery	1993	1	Melonycteris_fardoulsi	Flannery, T. F. (1993). Revision of the genus Melonycteris (Pteropodidae: Mammalia). Records of the Australian Museum, 45(1), 68.	https://journals.australian.museum/flannery-1993-rec-aust-mus-451-5980/	AM M.18833		"secondary forest near Sesena Village (8°31'S 162°05'E), altitude 100–200 m, Makira [= San Cristobal Island], Solomon Islands."	-8.51667	162.0833	fardoulisi (Flannery, 1993)	previously included N. maccoyi and N. mengermani; moved from Melonycteris to Nesonycteris	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.|Lavery, T. H., DeCicco, L. H., Olson, K. V., Holland, P. S., & Moyle, R. G. (2023). Phylogeography of Solomon Islands blossom bats reflects oceanic divides and Pleistocene connections. Journal of Biogeography, 00, 1-12.				Solomon Islands	Oceania	Australasia/Oceania	NT	0	0	0	Nesonycteris_fardoulisi	0	sciname match	Melonycteris_fardoulisi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Nesonycteris_fardoulisi	1004432	23	Fardoulis's Blossom Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Melonycterini	Nesonycteris	NA	fardoulisi	Flannery	1	Melonycteris fardoulisi	Flannery, T.F. 1993-03-19. Revision of the genus _Melonycteris_ (Pteropodidae: Mammalia). Records of the Australian Museum 45(1):59-80.	https://doi.org/10.3853/j.0067-1975.45.1993.130	AM M.18833	holotype		"secondary forest near Sesena Village (8°31'S 162°05'E), altitude 100–200 m, Makira [= San Cristobal Island], Solomon Islands."	-10.5167	162.0833	previously included N. maccoyi and N. mengermani; moved from Melonycteris to Nesonycteris	Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 2020(3950), 1-24.|Lavery, T. H., DeCicco, L. H., Olson, K. V., Holland, P. S., & Moyle, R. G. (2023). Phylogeography of Solomon Islands blossom bats reflects oceanic divides and Pleistocene connections. Journal of Biogeography, 00, 1-12.				Solomon Islands	Oceania (Continent)	Australasia	NT	0	0	0	Nesonycteris_fardoulisi	0	sciname match	Melonycteris_fardoulisi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Nesonycteris		fardoulisi	Flannery	1993	1	Rec. Aust. Mus.	46:08:00	Fardoulis' Blossom Bat	maccoyi Flannery, 1993; mengermani Flannery, 1993; schouteni Flannery, 1993.	Solomon Isls., Makira (= San Cristobal Isl.), Sesena	Makira (Solomon Isls)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13141/21978862/' target='_blank'>Near Threatened</a>	Previously considered a subgenus of Melonycteris (see e.g., Phillips, 1968; Flannery 1993; Simmons, 2005). Raised to full generic status by Almeida et al. (2020). Melonycteris, including Nesonycteris, was revised by Flannery (1993). Does not include maccoyi or mengermani; see Pulvers and Colgan (2006) and Lavery et al. (2023). See also Flannery (1995). Populations on Guadalcanal previously attributed to fardoulisi appear to belong to woodfordi; see Lavery et al. (2023). For phylogeography see Pulvers and Colgan (2006) and Lavery et al. (2023).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Melonycteris fardoulisi; Nesonycteris fardoulisi; Nesonycteris fardoulisi; Nesonycteris fardoulisi; Melonycteris fardoulisi; Nesonycteris fardoulisi; fardoulisi; maccoyi; mengermani; schouteni; fardoulisi; maccoyi; mengermani; schouteni; maccoyi; mengermani; schouteni; fardoulisi; maccoyi; mengermani; schouteni; Mélonyctére de Fardoulis; Fardoulis-Blitenflughund; Melonicterio de Fardoulis; Fardoulis's Blossom Bat; Fardoulis's Blossom Bat; Fardoulis' Blossom Bat; N. fardoulisi