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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1011	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti	Neopteryx frosti		[MSW3] Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).; [HMW] Neopteryx frosti Hayman, 1946 , “Tamalanti, West Celebes [= Sulawesi , Indonesia ], 3300 ft. [= 1006 m ].” Neopteryx is sister taxon to typical Pteropus and Acerodon . Monotypic.; [batnames2022] Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).; [batnames2023] Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).; [batnames2025_1.7] Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).														frosti				frosti 	frosti 			frosti Hayman, 1946		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Small-toothed fruit bat	W Celebes	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Neopteryx frosti	Indonesia, W. Sulawesi, Tamalanti, 3300 ft. (1006 m).	Hayman	1946	Ann. Mag. Nat. Hist., ser. 11, 12:571.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Small-toothed fruit bat	N, W Sulawesi	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hayman	1946	Ann. Mag. Nat. Hist., ser. 11, 12:571.		W and N Sulawesi.	Indonesia, W Sulawesi, Tamalanti, 3,300 ft. (1,006 m).		HAYMAN	1945	Size medium (fore arm length, 110 mm).	Distribution: Same as for genus.	No subspecies.		28	species	N. frosti	HAYMAN	1945	Neopteryx	genus	Neopteryx frosti				Size medium (fore arm length, 110 mm).	No subspecies.		1. N.frosti HAYMAN 1945.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Neopteryx frosti	Neopteryx		frosti	Hayman		1946		Ann. Mag. Nat. Hist., ser. 11	12		571		Small-toothed Fruit Bat	Indonesia, W Sulawesi, Tamalanti, 3,300 ft. (1,006 m).	W and N Sulawesi.	IUCN/SSC Action Plan (1992) – Rare: Limited Distribution. IUCN 2003 – Vulnerable.		Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).	03AD87FAFF8FF6668C7231E7FDDFFE67	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	133	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF8FF6668C7231E7FDDFFE67.xml	Neopteryx frosti	Pteropodidae	Neopteryx	frosti	Hayman	1946	Roussette de Frost @fr | Kleinzahnflughund @de | Nedptero de Frost @es	Neopteryx frosti Hayman, 1946 , “Tamalanti, West Celebes [= Sulawesi , Indonesia ], 3300 ft. [= 1006 m ].” Neopteryx is sister taxon to typical Pteropus and Acerodon . Monotypic.	Only known from two localities in C & NE Sulawesi (Talamanti and Bogani Nani Wartabone National Park).	Head-body c. 155 mm (tailless), ear 18- 6—20 mm , hindfoot with claws 30-5-33- 3 mm , forearm 104- 9- 111 mm ; weight 190-250 g . Males have shorter forearms than females but larger skull dimensions. Sepia-brown muzzle is crossed by three cream-white stripes: one stripe runs from tip of muzzle to forehead and other two along outside of muzzle to just behind end of eyes. Long woolly hair is present on upperparts of body, especially on back of mantle; individual hairs on mantle are brownish white dorsally and brownish only at tips, thereby giving clear demarcation from darker hairs on lower parts of back. Underparts of body have uniformly brown short fur, somewhat lighter at most at hair bases. Ventrally, collar area with its uniformly light brown hair is not as clearly separated. Thumb has well-developed claw; index claw is absent. Wing membrane begins at back near medial line and below between base offirst and second toe. Wing membranes are light to medium brown, with network pattern of blackish brown lines and spots. Rostrum is elongated and sharply tapering. Rami of mandible are thin. Posterior palatine foramina are large. Dental formulais12/2,C1/1,P 2/3, M 2/3 (x2) = 32. I! is slightly larger than I*. C! has narrow posterior basal shelf. Premolars and molars are small and simplified, with smooth and rounded crowns. P! is absent, and P? is higher but shorter than P*, M!, and M2.	Primary forests from lowlands to montane areas at elevations of 225-1006 m .	Narrowing ofpalate in front section, combined with tongue papillae and weakening ofteeth, suggest that the Small-toothed Fruit Bat eats rather soft fruit and flower products.	One Small-toothed Fruit Bat was pregnant with one embryo in March. Two other females bought at the market of Imandi were also pregnant in January and May. Embryos were 21-48- 5 mm . Male specimens have external penis of c. 12 mm length. Longevity is c.6 years.	The Small-toothed Fruit Bat is probably crepuscular or nocturnal. It presumably flies high in upper parts oftreetops. Two specimens were caught with mist nets high above the Sungei Tumpah River .	No information.	Classified as Endangered on The IUCN Red List. Despite intensive searches, only seven Small-toothed Fruit Bats have been described from two regions far away from each other. Captures have been limited to primary forests, which in Sulawesi are declining due to deforestation and spread of agriculture. Hunting for bushmeatis a threat, and hunters report thatit is becoming increasingly rare. Additional research and conservation programs are urgently needed. On following pages: 130. Sulawesi Flying Fox ( Acerodon celebensis ); 131. Talaud Flying Fox ( Acerodon humilis ); 132. Golden-capped Flying Fox ( Acerodon jubatus ); 133. Palawan Flying Fox ( Acerodon leucotis ): 134. Sunda Flying Fox ( Acerodon macklotil); 135. Moluccan Masked Flying Fox ( Pteropus personatus ); 136. Lombok Flying Fox ( Pteropus lombocensis ); 137. Little Red Flying Fox ( Pteropus scapulatus ); 138. Big-eared Flying Fox ( Pteropus macrotis ): 139. Gilliard's Flying Fox ( Pteropus gilliardorum ); 140. Sanborn’s Flying Fox ( Pteropus mahaganus ); 141. Dwarf Flying Fox ( Pteropus woodford); 142. Pohnpei Flying Fox ( Pteropus molossinus ); 143. Chuuk Flying Fox ( Pteropus pelagicus ).	Almeida et al. (2014) | Bergmans (2001) | Bergmans & Rozendaal (1988) | Hayman (1946) | Nowak (1999) | Pacifici et al. (2013) | Tsang (20164)		129. Small-toothed Fruit Bat Neopteryx frosti French: Roussette de Frost / German: Kleinzahnflughund / Spanish: Nedptero de Frost Taxonomy. Neopteryx frosti Hayman, 1946 , “Tamalanti, West Celebes [= Sulawesi , Indonesia ], 3300 ft. [= 1006 m ].” Neopteryx is sister taxon to typical Pteropus and Acerodon . Monotypic. Distribution. Only known from two localities in C & NE Sulawesi (Talamanti and Bogani Nani Wartabone National Park). Descriptive notes. Head-body c. 155 mm (tailless), ear 18- 6—20 mm , hindfoot with claws 30-5-33- 3 mm , forearm 104- 9- 111 mm ; weight 190-250 g . Males have shorter forearms than females but larger skull dimensions. Sepia-brown muzzle is crossed by three cream-white stripes: one stripe runs from tip of muzzle to forehead and other two along outside of muzzle to just behind end of eyes. Long woolly hair is present on upperparts of body, especially on back of mantle; individual hairs on mantle are brownish white dorsally and brownish only at tips, thereby giving clear demarcation from darker hairs on lower parts of back. Underparts of body have uniformly brown short fur, somewhat lighter at most at hair bases. Ventrally, collar area with its uniformly light brown hair is not as clearly separated. Thumb has well-developed claw; index claw is absent. Wing membrane begins at back near medial line and below between base offirst and second toe. Wing membranes are light to medium brown, with network pattern of blackish brown lines and spots. Rostrum is elongated and sharply tapering. Rami of mandible are thin. Posterior palatine foramina are large. Dental formulais12/2,C1/1,P 2/3, M 2/3 (x2) = 32. I! is slightly larger than I*. C! has narrow posterior basal shelf. Premolars and molars are small and simplified, with smooth and rounded crowns. P! is absent, and P? is higher but shorter than P*, M!, and M2. Habitat. Primary forests from lowlands to montane areas at elevations of 225-1006 m . Food and Feeding. Narrowing ofpalate in front section, combined with tongue papillae and weakening ofteeth, suggest that the Small-toothed Fruit Bat eats rather soft fruit and flower products. Breeding. One Small-toothed Fruit Bat was pregnant with one embryo in March. Two other females bought at the market of Imandi were also pregnant in January and May. Embryos were 21-48- 5 mm . Male specimens have external penis of c. 12 mm length. Longevity is c.6 years. Activity patterns. The Small-toothed Fruit Bat is probably crepuscular or nocturnal. It presumably flies high in upper parts oftreetops. Two specimens were caught with mist nets high above the Sungei Tumpah River . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Endangered on The IUCN Red List. Despite intensive searches, only seven Small-toothed Fruit Bats have been described from two regions far away from each other. Captures have been limited to primary forests, which in Sulawesi are declining due to deforestation and spread of agriculture. Hunting for bushmeatis a threat, and hunters report thatit is becoming increasingly rare. Additional research and conservation programs are urgently needed. On following pages: 130. Sulawesi Flying Fox ( Acerodon celebensis ); 131. Talaud Flying Fox ( Acerodon humilis ); 132. Golden-capped Flying Fox ( Acerodon jubatus ); 133. Palawan Flying Fox ( Acerodon leucotis ): 134. Sunda Flying Fox ( Acerodon macklotil); 135. Moluccan Masked Flying Fox ( Pteropus personatus ); 136. Lombok Flying Fox ( Pteropus lombocensis ); 137. Little Red Flying Fox ( Pteropus scapulatus ); 138. Big-eared Flying Fox ( Pteropus macrotis ): 139. Gilliard's Flying Fox ( Pteropus gilliardorum ); 140. Sanborn’s Flying Fox ( Pteropus mahaganus ); 141. Dwarf Flying Fox ( Pteropus woodford); 142. Pohnpei Flying Fox ( Pteropus molossinus ); 143. Chuuk Flying Fox ( Pteropus pelagicus ). Bibliography. Almeida et al. (2014), Bergmans (2001), Bergmans & Rozendaal (1988), Hayman (1946), Nowak (1999), Pacifici et al. (2013), Tsang (20164).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Neopteryx frosti	Neopteryx		frosti	Hayman	1946	0	Ann. Mag. Nat. Hist.	ser. 11, 12: 571	Small-toothed Fruit Bat	None.	Indonesia, W Sulawesi, Tamalanti, 3,300 ft. (1,006 m).	W and N Sulawesi.	Not listed.	Endangered	Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Neopteryx frosti	23	Small-toothed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTEROPODINI	Neopteryx	NA	frosti	Hayman	1946	0						"Tamalanti, West Celebes [= Sulawesi, Indonesia], 3300 ft. [= 1006 m]."			frosti Hayman, 1946	NA	NA	Indonesia	Asia	Australasia/Oceania	EN	0	0	0	Neopteryx_frosti	0	sciname match	Neopteryx_frosti	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14560	Neopteryx frosti	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Neopteryx	frosti	Hayman, 1946		100000000	Neopteryx frosti	Endangered	C1	2016	2016-06-01 00:00:00 UTC	3.1	English	Neopteryx frosti is assessed as Endangered under criterion C1. The species is estimated to have fewer than 2,500 mature individuals and is projected to undergo a 20% reduction over two generations (12 years as per Pacifici et al.  2013) due to continued habitat decline and intense hunting. It is known from only two widely separated localities and intensive survey work has not recorded this species else where, and both localities are within primary forest, which is declining rapidly on Sulawesi. The species is also present in bushmeat markets, though hunters have reported that they are increasingly rare.	It inhabits primary forest. Two specimens were mist-netted high over a river but there is no other information known.	Deforestation due to logging and expanding agriculture is a major threat, as the species prefers to roost in primary forests. Hunting of what appears to be a rare and restricted range species is also a threat, four of the seven known specimens are from markets.	This is a rare species, known from only seven specimens (Bergmans and Rozendaal 1988, Bergmans 2001). Extensive mist-netting efforts have failed to find the species, although appropriate survey methods may not have been used (the only mist-netted specimens came from above the canopy). Hunters interviewed in bushmeat markets have expressed that this species has become even rarer in recent years, and none were found from recent market surveys (Sheherazade, pers. comm).	Decreasing	This fruit bat is known only from two localities on Sulawesi, Indonesia at 225 and 1,000 m asl, despite extensive surveying. The species has not been caught in any of these localities since 1991. It is possibly extinct from Central Sulawesi, as that area has been subject to a the high level of habitat disturbance and logging.	The species is present in bushmeat markets.	Terrestrial	The species has been recorded from Bogani Nani Wartabone National Park.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Neopteryx		frosti	Hayman	1946	0	Ann. Mag. Nat. Hist.	ser. 11, 12: 571	Small-toothed Fruit Bat	None.	Indonesia, W Sulawesi, Tamalanti, 3,300 ft. (1,006 m).	W and N Sulawesi.	Not listed.	Endangered	Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).	Neopteryx frosti	1004447	23	Small-toothed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTEROPODINI	Neopteryx	NA	frosti	Hayman	1946	0						"Tamalanti, West Celebes [= Sulawesi, Indonesia], 3300 ft. [= 1006 m]."			frosti Hayman, 1946	NA	NA				Indonesia	Asia	Australasia/Oceania	EN	0	0	0	Neopteryx_frosti	0	sciname match	Neopteryx_frosti	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Neopteryx_frosti	1004447	23	Small-toothed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteropodini	Neopteryx	NA	frosti	Hayman	0	Neopteryx frosti	Hayman, R.W. 1946-07-10. A new genus of fruit-bat and a new squirrel, from Celebes. Annals and Magazine of Natural History (11)12(93):569-578.	https://doi.org/10.1080/00222934508654763	BMNH:Mamm:1940.691k	holotype	https://data.nhm.ac.uk/object/3417a1e1-16d9-470d-a908-0ae0041455d3	"Tamalanti, West Celebes [= Sulawesi, Indonesia], 3300 ft. [= 1006 m]."			NA	NA				Indonesia	Asia	Australasia	EN	0	0	0	Neopteryx_frosti	0	sciname match	Neopteryx_frosti	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Neopteryx		frosti	Hayman	1946	0	Ann. Mag. Nat. Hist.	ser. 11, 12: 571	Small-toothed Fruit Bat	None.	Indonesia, W Sulawesi, Tamalanti, 3,300 ft. (1,006 m).	W and N Sulawesi.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14560/115122474/' target='_blank'>Endangered</a>	Known from only 7 specimens; see Bergmans and Rozendaal (1988) and Bergmans (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Neopteryx frosti; Neopteryx frosti; Neopteryx frosti; Neopteryx frosti; Neopteryx frosti; Neopteryx frosti; frosti; Roussette de Frost; Kleinzahnflughund; Nedptero de Frost; Small-toothed Fruit Bat; Small-toothed Fruit Bat; Small-toothed Fruit Bat; N. frosti