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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L1002	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Eptesicus diminutus	Neoeptesicus diminutus	Neoeptesicus diminutus	Neoeptesicus diminutus		[MSW2] Subgenus Eptesicus. Includes fidelis; see Williams (1978c:380-382), and also for the use of diminutus instead of dorianus.; [MSW3] Subgenus Eptesicus. Includes fidelis but does not include dorianus; see Williams (1978c).; [HMW] Eptesicus diminutus Osgood, 1915 , “Sao Marcello, Rio Preto, Bahia , Brazil .” G. E. Dobson in 1885 considered E. diminutus a synonym of E. dorianus, but qualitative and quantitative morphological traits of the holotype most resemble E. brasiliensis . Moreover, there is no certainty about the origin of this holotype, or if it could be changed with other individuals at some point. Therefore, the name E. dorianus was considered a nomen dubium (dubious name), and the next available name, E. diminutus , was applied to the taxon. An isolated population in Venezuela is probably a new subspecies or species. Two subspecies recognized.; [batnames2022] Subgenus Eptesicus .  Includes fidelis but does not include dorianus ; see Williams (1978 c ).; [batnames2023] Subgenus Eptesicus .  Includes fidelis but does not include dorianus ; see Williams (1978 c ).; [MDD2025_2.0] moved from Eptesicus to the recently described Neoeptesicus; [batnames2025_1.7] Includes fidelis but does not include dorianus; see Williams (1978c).; [MDD2025_2.2] moved from Eptesicus to the recently described Neoeptesicus				fidelis, dorianus		fidelis.	diminutus, fidelis	diminutus, fidelis		diminutus, fidelis		diminutus, fidelis		diminutus, fidelis		diminutus, fidelis		diminutus, fidelis	diminutus, fidelis, dimidiatus, fidens	diminutus, fidelis		diminutus (Osgood, 1915)|fidelis (O. Thomas, 1920)|dimidiatus (Handley, 1976) [incorrect subsequent spelling]|fidens (J. Muñoz, 2001) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		E, SE Brazil – N Argentina, Uruguay; ref. 4.88	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Eptesicus diminutus	Brazil, Bahia, Rio Preto, Sao Marcello.	Osgood	1915	Field Mus. Nat. Hist. Publ. Ser. Zool., 10:197.	Distribution: Ranging from eastern Brazil to northern Argentina.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		E, SE Brazil – N Argentina, Uruguay	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Osgood	1915	Field Mus. Nat. Hist. Publ., Zool. Ser., 10:197.	Subgenus Eptesicus. Includes fidelis; see Williams (1978c:380-382), and also for the use of diminutus instead of dorianus.	Venezuela, E Brazil, Paraguay, Uruguay, N Argentina.	Brazil, Bahia, Rio Preto, Sao Marcello.		OSGOOD	1915	Rostrum of medium width. Braincase fairly high. Size fairly small (forearm length, 31-38 mm). Fur relatively dark in color.	Distribution: Ranging from eastern Brazil to northern Argentina.	Two subspecies are currently recognized:	E. d. diminutus (east-central Brazil), E. d. fidelis (extreme southeastern Brazil to northern Argentina).	121	species	E. diminutus	OSGOOD	1915	Eptesicus	subgenus	Eptesicus diminutus				Rostrum of medium width. Braincase fairly high. Size fairly small (forearm length, 31-38 mm). Fur relatively dark in color.	Two subspecies are currently recognized:		15. E. diminutus OSGOOD 1915 [innoxius group],	15	_N. d. diminutus_ (Osgood, 1915); _N. d. fidelis_ (Thomas, 1920)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Eptesicini	Eptesicus diminutus	Eptesicus	Eptesicus	diminutus	Osgood		1915		Field Mus. Nat. Hist. Publ., Zool. Ser.	10		197		Diminutive Serotine	Brazil, Bahia, Rio Preto, São Marcello.	Venezuela, E Brazil, Paraguay, Uruguay, N Argentina.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	fidelis Thomas, 1920.	Subgenus Eptesicus. Includes fidelis but does not include dorianus; see Williams (1978c).	4C3D87E8FFAD6A12FF449168192DB80B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	844	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFAD6A12FF449168192DB80B.xml	Eptesicus diminutus	Vespertilionidae	Eptesicus	diminutus	Osgood	1915	Petite Sérotine @fr | Kleine Breitflligelfledermaus @de | Eptesicus pequeno @es	Eptesicus diminutus Osgood, 1915 , “Sao Marcello, Rio Preto, Bahia , Brazil .” G. E. Dobson in 1885 considered E. diminutus a synonym of E. dorianus, but qualitative and quantitative morphological traits of the holotype most resemble E. brasiliensis . Moreover, there is no certainty about the origin of this holotype, or if it could be changed with other individuals at some point. Therefore, the name E. dorianus was considered a nomen dubium (dubious name), and the next available name, E. diminutus , was applied to the taxon. An isolated population in Venezuela is probably a new subspecies or species. Two subspecies recognized.	E.d.diminutusOsgood,1915—NVenezuelaandEBrazil. E. d. fidelis Thomas, 1920 — S Brazil , Paraguay , Uruguay , and N & C Argentina .	Head-body ¢.45-63 mm,tail 33-42 mm, ear 11-13-6 mm, hindfoot 6-10 mm, forearm 30-37-8 mm; weight 4-5-7 g. Females are larger than males. Dorsal hairs of the Diminutive Serotine are bicolored, with dark brown bases and brown to yellowish brown tips; sometimes extreme distal parts of hairs can be much paler, giving slightly frosted appearance. Ventral hairs have dark brown bases and pale brown to whitish tips. Some populations in Argentina have generally brown to dark brown fur, with almost unicolored hairs and slightly lighter venter. Ears are medium-sized, with rounded tips, and are well separated; tragusis relatively broad and short. Membranes are dark grayish to blackish. Skull is short and slender; sagittal crest is poorly developed but visible; lambdoidal crests are visible; zygomatic arches are thin, strong, and slightly widened medially; pterygoids are well developed; and tympanic bullae are large. Upper inner incisors are separated, bilobed, and spatulated; I’ is reduced, conical, and separated from C' by small gap; P* is well developed, reaching one-third of C' height; M' and M? are almost square, with W-pattern; M” is reduced and triangular; lower incisors are trilobed and in contact,filling all space between canines; P, is small, reaching onethird of P, height; and lower molars have well-developed cusps, decreasing in size from M, to M,. Chromosomal complement has 2n = 50 and FN = 48, with acrocentric autosomes, a large submetacentric X-chromosome, and a small acrocentric Y-chromosome.	Dry tropical and subtropical forests in primary and secondary forests, open areas, grasslands, dense forests, and urban areas from sea level up to elevations of c. 1157 m . Diminutive Serotines have been caught on forest edges, in clustered environments, and over streams.	The Diminutive Serotine is insectivorous. Stomach contents had fragments of Coleoptera , Lepidoptera , and Diptera .	Lactating Diminutive Serotines and a juvenile were captured in late January in southern Brazil .	Diminutive Serotines roost under loose bark, hollow trees, and buildings. Echolocation calls have high frequencies of ¢.80 kHz and low frequencies of c.40 kHz. Calls are c.1-8 milliseconds in duration, mean interpulse interval is 50 milliseconds, and frequency of maximum energy is c.66 kHz.	No information.	Classified as Least Concern on The IUCNRed List. The Diminutive Serotine is widely distributed and found in protected areas, but it is not locally common.	Arias-Aguilar et al. (2018) | Barquez et al. (1999) | Bianconi & Pedro (2017) | Davis, W.B. (1966) | Davis, W.B. & Gardner (2008) | Dobson (1885) | Gonzalez & Barquez (2016a) | Myers et al. (1983) | Simmons (2005) | Solari et al. (2013) | Williams (1978)	https://zenodo.org/record/6398188/files/figure.png	180. Diminutive Serotine Eptesicus diminutus French: Petite Sérotine / German: Kleine Breitflligelfledermaus / Spanish: Eptesicus pequeno Taxonomy. Eptesicus diminutus Osgood, 1915 , “Sao Marcello, Rio Preto, Bahia , Brazil .” G. E. Dobson in 1885 considered E. diminutus a synonym of E. dorianus, but qualitative and quantitative morphological traits of the holotype most resemble E. brasiliensis . Moreover, there is no certainty about the origin of this holotype, or if it could be changed with other individuals at some point. Therefore, the name E. dorianus was considered a nomen dubium (dubious name), and the next available name, E. diminutus , was applied to the taxon. An isolated population in Venezuela is probably a new subspecies or species. Two subspecies recognized. Subspecies and Distribution. E.d.diminutusOsgood,1915—NVenezuelaandEBrazil. E. d. fidelis Thomas, 1920 — S Brazil , Paraguay , Uruguay , and N & C Argentina . Descriptive notes. Head-body ¢.45-63 mm,tail 33-42 mm, ear 11-13-6 mm, hindfoot 6-10 mm, forearm 30-37-8 mm; weight 4-5-7 g. Females are larger than males. Dorsal hairs of the Diminutive Serotine are bicolored, with dark brown bases and brown to yellowish brown tips; sometimes extreme distal parts of hairs can be much paler, giving slightly frosted appearance. Ventral hairs have dark brown bases and pale brown to whitish tips. Some populations in Argentina have generally brown to dark brown fur, with almost unicolored hairs and slightly lighter venter. Ears are medium-sized, with rounded tips, and are well separated; tragusis relatively broad and short. Membranes are dark grayish to blackish. Skull is short and slender; sagittal crest is poorly developed but visible; lambdoidal crests are visible; zygomatic arches are thin, strong, and slightly widened medially; pterygoids are well developed; and tympanic bullae are large. Upper inner incisors are separated, bilobed, and spatulated; I’ is reduced, conical, and separated from C' by small gap; P* is well developed, reaching one-third of C' height; M' and M? are almost square, with W-pattern; M” is reduced and triangular; lower incisors are trilobed and in contact,filling all space between canines; P, is small, reaching onethird of P, height; and lower molars have well-developed cusps, decreasing in size from M, to M,. Chromosomal complement has 2n = 50 and FN = 48, with acrocentric autosomes, a large submetacentric X-chromosome, and a small acrocentric Y-chromosome. Habitat. Dry tropical and subtropical forests in primary and secondary forests, open areas, grasslands, dense forests, and urban areas from sea level up to elevations of c. 1157 m . Diminutive Serotines have been caught on forest edges, in clustered environments, and over streams. Food and Feeding. The Diminutive Serotine is insectivorous. Stomach contents had fragments of Coleoptera , Lepidoptera , and Diptera . Breeding. Lactating Diminutive Serotines and a juvenile were captured in late January in southern Brazil . Activity patterns. Diminutive Serotines roost under loose bark, hollow trees, and buildings. Echolocation calls have high frequencies of ¢.80 kHz and low frequencies of c.40 kHz. Calls are c.1-8 milliseconds in duration, mean interpulse interval is 50 milliseconds, and frequency of maximum energy is c.66 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCNRed List. The Diminutive Serotine is widely distributed and found in protected areas, but it is not locally common. Bibliography. Arias-Aguilar et al. (2018), Barquez et al. (1999), Bianconi & Pedro (2017), Davis, W.B. (1966), Davis, W.B. & Gardner (2008), Dobson (1885), Gonzalez & Barquez (2016a), Myers et al. (1983), Simmons (2005), Solari et al. (2013), Williams (1978).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus diminutus	Eptesicus	Eptesicus	diminutus	Osgood	1915	0	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.5535	Diminutive Serotine	<b> fidelis </b> Thomas, 1920.	Brazil, Bahia, Rio Preto, S&atilde;oMarcello.	Venezuela, E Brazil, Paraguay, Uruguay, N Argentina.	Not listed.	Least Concern	Subgenus Eptesicus .  Includes fidelis but does not include dorianus ; see Williams (1978 c ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Eptesicus diminutus	23	Diminutive Serotine	Little Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Eptesicus	Eptesicus	diminutus	Osgood	1915	0	Eptesicus_diminutus	Osgood, W. H. (1915). New mammals from Brazil and Peru. Field Museum of Natural History Zoological Series, 10, 197.	https://www.biodiversitylibrary.org/item/20379#page/15/mode/1up	FMNH 20971		"Saõ Marcello, Rio Preto, Bahia, Brazil."			diminutus Osgood, 1915|fidelis O. Thomas, 1920	NA	NA	Venezuela|Brazil|Paraguay|Uruguay|Argentina	South America	Neotropic	LC	0	0	0	Eptesicus_diminutus	0	sciname match	Eptesicus_diminutus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	7922	Eptesicus diminutus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Eptesicus	diminutus	Osgood, 1915		100000000	Eptesicus diminutus	Least Concern		2016	2016-06-24 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution; although it is not a common bat, it occurs at several protected areas of different scale and nature. The northern population (Venezuela) could represent an undescribed subspecies and therefore require an individual assessment in view of its isolation.	The species is insectivorous, it inhabits open areas, and urban and sub-urban areas with low populations (Barquez et al. 1999).	This species is not threatened.	The species is not common, but has a wide distribution.	Unknown	Eptesicus diminutus occurs through Venezuela, Brazil, Paraguay, Uruguay, and northern Argentina (Simmons 2005, Gardner 2008). The species has not been recorded in Colombia (Solari et al. 2013).		Terrestrial	Further studies are needed to check taxonomic status of the species, and the systematics of diminuta's group.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Eptesicus	Eptesicus	diminutus	Osgood	1915	0	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.553472	Diminutive Serotine	<b> fidelis </b> Thomas, 1920.	Brazil, Bahia, Rio Preto, S&atilde;oMarcello.	Venezuela, E Brazil, Paraguay, Uruguay, N Argentina.	Not listed.	Least Concern	Subgenus Eptesicus .  Includes fidelis but does not include dorianus ; see Williams (1978 c ).	Eptesicus diminutus	1005516	23	Diminutive Serotine	Little Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Eptesicus	Eptesicus	diminutus	Osgood	1915	0	Eptesicus_diminutus	Osgood, W. H. (1915). New mammals from Brazil and Peru. Field Museum of Natural History Zoological Series, 10, 197.	https://www.biodiversitylibrary.org/item/20379#page/15/mode/1up	FMNH 20971		"Saõ Marcello, Rio Preto, Bahia, Brazil."			diminutus Osgood, 1915|fidelis O. Thomas, 1920	NA	NA				Venezuela|Brazil|Paraguay|Uruguay|Argentina	South America	Neotropic	LC	0	0	0	Eptesicus_diminutus	0	sciname match	Eptesicus_diminutus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Neoeptesicus_diminutus	1005516	23	Diminutive Serotine	Little Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Neoeptesicus	NA	diminutus	Osgood	1	Eptesicus diminutus	Osgood, W.H. 1915-10-22. New mammals from Brazil and Peru. Field Museum of Natural History, Zoological Series 10(13):187-198.	https://www.biodiversitylibrary.org/page/2643065	FMNH:Mamm:20971	holotype	http://portal.vertnet.org/o/fmnh/mammals?id=9d613910-4c1c-4d1e-ad54-0613f14836d9	"Saõ Marcello, Rio Preto, Bahia, Brazil."			moved from Eptesicus to the recently described Neoeptesicus	Cláudio, V. C., Novaes, R. L., Gardner, A. L., Nogueira, M. R., Wilson, D. E., Maldonado, J. E., ... & Moratelli, R. (2023). Taxonomic re-evaluation of New World Eptesicus and Histiotus (Chiroptera: Vespertilionidae), with the description of a new genus. Zoologia (Curitiba), 40, e22029.				Venezuela|Brazil|Paraguay|Uruguay|Argentina|Bolivia	South America	Neotropic	LC (as Eptesicus diminutus)	0	0	0	Eptesicus_diminutus	0	sciname match	Eptesicus_diminutus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Neoeptesicus		diminutus	Osgood	1915	1	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.553472	Diminutive Serotine	fidelis Thomas, 1920.	Brazil, Bahia, Rio Preto, S&atilde;oMarcello.	Venezuela, E Brazil, Paraguay, Uruguay, N Argentina.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/7922/115087028/' target='_blank'>Least Concern as Eptesicus diminutus</a>	Includes fidelis but does not include dorianus; see Williams (1978c).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Eptesicus diminutus; Eptesicus diminutus; Eptesicus diminutus; Eptesicus diminutus; Eptesicus diminutus; Eptesicus diminutus; diminutus; fidelis; diminutus; fidelis; fidelis; diminutus; fidelis; Petite Sérotine; Kleine Breitflligelfledermaus; Eptesicus pequeno; Diminutive Serotine; Little Serotine; Diminutive Serotine; Diminutive Serotine; E. diminutus