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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L101	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Barbastella leucomelas [synonym of]	Barbastella leucomeles darjelingensis	Barbastella leucomeles darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis	Barbastella darjelingensis		[HMW] Plecotus darjelingensis Hodgson in Horsfield, 1855 , Darjeeling, India . Previously treated as a race of B. leucomelas ; separated, based mainly on genetic and morphological data. This species included B. pacifica until the very recent taxonomic review by S. V. Kruskop and colleagues in 2019, when latter was recognized as a full species. Chinese and South-east Asian representatives of B. darjelingensis may represent a furtherspecies, distinct from the Himalayan true B. darjelingensis ; indeed, B. pacifica is genetically sister to the Chinese and South-east Asian B. darjelingensis , and together they are sister to the rest of Barbastella . Further study needed. Monotypic.; [batnames2022] Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).; [MDD2022] split from B. leucomeles; [IUCN] Barbastella darjelingensis ; was previously considered a subspecies to ;B. leucomelas ;(as ;B. l. darjelingensis ), however, ;B. leucomelas ;s.str. was shown to be a separate species by Benda ;et al. ;(2008). There is a variety of mitochondrial lineages within this species (Kruskop ;et al. ; 2019, Chakravarty ;et al. ;2020), suggesting the possible presence of cryptic taxa. However, at least the mainland specimens are similar in appearance and do not differ in craniometry (Kruskop ;et al. ;2019), therefore, until convincing evidence to the contrary, all ;Barbastella populations from eastern Afghanistan to Taiwan could be considered as a one species,Barbastella darjelingensis . This does not include populations from Central Asia (Kruskop 2015) and Japan (Kruskop ;et al. ;2019), which represent separate species.; [batnames2023] Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).; [MDD2023] split from B. leucomeles; [MDD2025_2.0] split from B. leucomeles; [batnames2025_1.7] Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).; [MDD2025_2.2] split from B. leucomeles														darjelingensis, dargelinensis, blanfordi	Barbastella darjelingensis ; was previously considered a subspecies to ;B. leucomelas ;(as ;B. l. darjelingensis ), however, ;B. leucomelas ;s.str. was shown to be a separate species by Benda ;et al. ;(2008). There is a variety of mitochondrial lineages within this species (Kruskop ;et al. ; 2019, Chakravarty ;et al. ;2020), suggesting the possible presence of cryptic taxa. However, at least the mainland specimens are similar in appearance and do not differ in craniometry (Kruskop ;et al. ;2019), therefore, until convincing evidence to the contrary, all ;Barbastella populations from eastern Afghanistan to Taiwan could be considered as a one species,Barbastella darjelingensis . This does not include populations from Central Asia (Kruskop 2015) and Japan (Kruskop ;et al. ;2019), which represent separate species.			darjelingensis, dargelinensis, blanfordi	darjelingensis, darjilingensis, dargelinensis, darjelinensis, blanfordi, formosanus			darjelingensis (B. H. Hodgson in Horsfield, 1855)|darjilingensis (J. E. Gray, 1863) [incorrect subsequent spelling]|dargelinensis Dobson, 1875 [incorrect subsequent spelling]|darjelinensis (Dobson, 1876) [incorrect subsequent spelling]|blanfordi Bianchi, 1917|formosanus Lin Liangkong, H.-C. Zheng, & Maeda, 1997 [nomen nudum]						N/A																																								NA																											4C3D87E8FF9C6A2CFA8F9B521A9FB345	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	861	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF9C6A2CFA8F9B521A9FB345.xml	Barbastella darjelingensis	Vespertilionidae	Barbastella	darjelingensis		1855	Barbastelle d'Asie @fr | Ostliche Mopsfledermaus @de | Barbastelaasiética @es | Asian Barbastelle @en	Plecotus darjelingensis Hodgson in Horsfield, 1855 , Darjeeling, India . Previously treated as a race of B. leucomelas ; separated, based mainly on genetic and morphological data. This species included B. pacifica until the very recent taxonomic review by S. V. Kruskop and colleagues in 2019, when latter was recognized as a full species. Chinese and South-east Asian representatives of B. darjelingensis may represent a furtherspecies, distinct from the Himalayan true B. darjelingensis ; indeed, B. pacifica is genetically sister to the Chinese and South-east Asian B. darjelingensis , and together they are sister to the rest of Barbastella . Further study needed. Monotypic.	N Afghanistan E through Himalayas to C & E China , N Laos , and N Vietnam , including Taiwan .	Head-body 47-51 mm , tail 40-47 mm , ear 15-17 mm , hindfoot 7-8 mm , forearm 38.7-42- 1 mm ; weight c. 9 g . Very similar to congeners. Fur is c. 8— 9 mm long on dorsum and 7-8 mm on belly. All taxonomically confirmed specimens have dark fur with frosted silver hair tips. Face is similar to other barbastelles, with short snout and wide squarish, broad, short ears, which join across forehead; tragus generally pointed, one-half ear height, without antitragal lobe; ears do not have typical rounded or circular lobe on outer margin of some other barbastelles. Muzzle short and broad. Wings relatively wide, inserting at base of first toe. Baculum is similar in size, but narrower than in the Arabian Barbastelle ( B. leucomelas ), parallel-sided, and not gradually tapering to epiphysis. Anterior one-half of braincase is bulbous; condylobasal length is c. 14 mm . Chromosomal complement has 2n = 32 and FN = 50.	Seems to select positively dry coniferous woodlands as its preferred foraging and roosting habitats. In Taiwan , it has been found in oak forests dominated by broadleaved trees, as well as plantations and pine woodland, usually in montane areas. Found from sea level up to 2500 m , including Himalayan moist temperate forests.	Feeds mainly on moths, using swallow-like flight.	Maternity colonies are thought to be small, with up to ¢.30 mature females. Reported to give birth to twins.	Roosts in caves, mines, tunnels, rock crevices, or hollow trees, or under bark.	Thought to be solitary. Males and females seem to roost separately, especially during the breeding season.	Not assessed on The IUCN Red List. The Eastern Barbastelle 1s included under the Arabian Barbastelle, which 1s classified as Least Concern. Deforestation and habitat loss might be a potential threat to this species, but threats need to be reassessed after recent taxonomic review.	Abe et al. (2005) | Acharya & Ruedas (2007) | Bates & Harrison (1997) | Benda & Mlikovsky (2008) | Das (2003) | Dietz & Kiefer (2016) | Duckworth & Pons (2011) | Horaéek et al. (2000) | Hutson et al. (2001) | Kruskop & Shchinov (2010) | Kruskop, Borisenko et al. (2012) | Kruskop, Kawai & Tiunov (2019) | Lin Liangkong, Lee Lingling & Cheng Hsichi (1997) | Lin Liangkong, Motokawa, Harada & Cheng Hsichi (2002) | Monadjem, Tsytsulina et al. (2017) | Osborn (1963) | Pacifici et al. (2013) | Shrestha (1997) | Simmons (2005) | Smith & Xie Yan (2008) | Uchida & Ando (1972) | Wang Yingxiang (2003) | Yoshiyuki (1989) | Zhang Jinshuo et al. (2007)	https://zenodo.org/record/6398279/files/figure.png	222. Eastern Barbastelle Barbastella darjelingensis French: Barbastelle d'Asie / German: Ostliche Mopsfledermaus / Spanish: Barbastela asiética Other common names: Asian Barbastelle Taxonomy. Plecotus darjelingensis Hodgson in Horsfield, 1855 , Darjeeling, India . Previously treated as a race of B. leucomelas ; separated, based mainly on genetic and morphological data. This species included B. pacifica until the very recent taxonomic review by S. V. Kruskop and colleagues in 2019, when latter was recognized as a full species. Chinese and South-east Asian representatives of B. darjelingensis may represent a furtherspecies, distinct from the Himalayan true B. darjelingensis ; indeed, B. pacifica is genetically sister to the Chinese and South-east Asian B. darjelingensis , and together they are sister to the rest of Barbastella . Further study needed. Monotypic. Distribution. N Afghanistan E through Himalayas to C & E China , N Laos , and N Vietnam , including Taiwan . Descriptive notes. Head-body 47-51 mm , tail 40-47 mm , ear 15-17 mm , hindfoot 7-8 mm , forearm 38.7-42- 1 mm ; weight c. 9 g . Very similar to congeners. Fur is c. 8— 9 mm long on dorsum and 7-8 mm on belly. All taxonomically confirmed specimens have dark fur with frosted silver hair tips. Face is similar to other barbastelles, with short snout and wide squarish, broad, short ears, which join across forehead; tragus generally pointed, one-half ear height, without antitragal lobe; ears do not have typical rounded or circular lobe on outer margin of some other barbastelles. Muzzle short and broad. Wings relatively wide, inserting at base of first toe. Baculum is similar in size, but narrower than in the Arabian Barbastelle ( B. leucomelas ), parallel-sided, and not gradually tapering to epiphysis. Anterior one-half of braincase is bulbous; condylobasal length is c. 14 mm . Chromosomal complement has 2n = 32 and FN = 50. Habitat. Seems to select positively dry coniferous woodlands as its preferred foraging and roosting habitats. In Taiwan , it has been found in oak forests dominated by broadleaved trees, as well as plantations and pine woodland, usually in montane areas. Found from sea level up to 2500 m , including Himalayan moist temperate forests. Food and Feeding. Feeds mainly on moths, using swallow-like flight. Breeding. Maternity colonies are thought to be small, with up to ¢.30 mature females. Reported to give birth to twins. Activity patterns. Roosts in caves, mines, tunnels, rock crevices, or hollow trees, or under bark. Movements, Home range and Social organization. Thought to be solitary. Males and females seem to roost separately, especially during the breeding season. Status and Conservation. Not assessed on The IUCN Red List. The Eastern Barbastelle 1s included under the Arabian Barbastelle, which 1s classified as Least Concern. Deforestation and habitat loss might be a potential threat to this species, but threats need to be reassessed after recent taxonomic review. Bibliography. Abe et al. (2005), Acharya & Ruedas (2007), Bates & Harrison (1997), Benda & Mlikovsky (2008), Das (2003), Dietz & Kiefer (2016), Duckworth & Pons (2011), Horaéek et al. (2000), Hutson et al. (2001), Kruskop & Shchinov (2010), Kruskop, Borisenko et al. (2012), Kruskop, Kawai & Tiunov (2019), Lin Liangkong, Lee Lingling & Cheng Hsichi (1997), Lin Liangkong, Motokawa, Harada & Cheng Hsichi (2002), Monadjem, Tsytsulina et al. (2017), Osborn (1963), Pacifici et al. (2013), Shrestha (1997), Simmons (2005), Smith & Xie Yan (2008), Uchida & Ando (1972), Wang Yingxiang (2003), Yoshiyuki (1989), Zhang Jinshuo et al. (2007).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Barbastella darjelingensis	Barbastella		darjelingensis	Hodgson [ in Horsfeld]	1855	0	Ann. Mag. Nat. Hist.	ser. 2, 16: 102	Darjeeling Barbastelle	None.		Taiwan; Sichuan China, N India, Nepal	Not listed.	Least Concern	Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Barbastella darjelingensis	23	Eastern Barbastelle	Asian Barbastelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PLECOTINI	Barbastella	NA	darjelingensis	Hodgson	1855	1	Plecotus_darjelingensis	Hodgson, B. H. (1855). In Horsfield, T. Brief notices of several new or little-known species of mammalia, lately discovered and collected in Nepal, by Brian Houghton Hodgson, Esq. Annals and Magazine of Natural History, Ser. 2, 16, 103.	https://www.biodiversitylibrary.org/page/2263778#page/130/mode/1up	BM 1854.9.1.13		Darjeeling, India.			darjelingensis (Hodgson, 1855)|dargelinensis Dobson, 1875|blanfordi Bianchi, 1917	split from B. leucomeles	Zhang, J. S., Han, N. J., Jones, G., Lin, L. K., Zhang, J. P., Zhu, G. J., ... & Zhang, S. Y. (2007). A new species of Barbastella (Chiroptera: Vespertilionidae) from north China. Journal of Mammalogy, 88(6), 1393-1403.	Afghanistan|Pakistan|India|Nepal|Bhutan|Bangladesh|Myanmar|Laos|Vietnam|China|Taiwan	Asia	Palearctic	LC	0	0	0	Barbastella_darjelingensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Barbastella darjelingensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Barbastella	darjelingensis	(Hodgson, 1855)	Barbastella darjelingensis ; was previously considered a subspecies to ;B. leucomelas ;(as ;B. l. darjelingensis ), however, ;B. leucomelas ;s.str. was shown to be a separate species by Benda ;et al. ;(2008). There is a variety of mitochondrial lineages within this species (Kruskop ;et al. ; 2019, Chakravarty ;et al. ;2020), suggesting the possible presence of cryptic taxa. However, at least the mainland specimens are similar in appearance and do not differ in craniometry (Kruskop ;et al. ;2019), therefore, until convincing evidence to the contrary, all ;Barbastella populations from eastern Afghanistan to Taiwan could be considered as a one species,Barbastella darjelingensis . This does not include populations from Central Asia (Kruskop 2015) and Japan (Kruskop ;et al. ;2019), which represent separate species.	90000000	Barbastella darjelingensis	Least Concern		2021	2020-12-10 00:00:00 UTC	3.1	English	Barbastella darjelingensis is assessed as Least Concern as it has a wide range, part of which is located in sparsely populated regions, so it is difficult to assume threats to its existence in the near future. However, careful research is needed over most of the range, as the species may include cryptic taxa, which may have a higher threat status.	Strictly nocturnal species. Little is known about its habits, but, by analogy with other barbastelles, it is probably solitary or forming small colonies and feeding on middle-size flying insects. Known mainly from mountain areas up to elevation of ca. 2,350 m ASL. Inhabiting mountainous temperate and subtropical forests. In Vietnam it was found foraging over stream in a deciduous forest at the elevation of ca. 2000 m ASL (Kruskop and Shchinov 2010), in India animals were captured in an oak forest and over a forest clearing (Chakravarty et al. ;2020). Day roosts probably in natural and artificial caves and caverns, rock crevices or in hollows under tree bark.	This species can be threatened by uncontrolled visits and development of underground cavities that serve as roosts for it, as well as the destruction of habitats and the use of pesticides in the surrounding agricultural lands.	Considering its wide range there seems to be little information available about the population of this species. According to published records, this species is highly sporadic, with low density of population, however it could be an artifact caused by distribution in hard-to-reach areas. The species population is presumed stable.	Stable	As accepted here, the species has a vast distribution range from eastern Afghanistan (Benda and Gaisler 2015) through northern India, Nepal, Bhutan, south China, to Laos, northern Vietnam and Taiwan (Bates and Harrison 1997, Smith and Xie 2008, Kruskop 2013). Known distribution connected with the mountainous areas of moderate elevations.	This species has no commercial value and not known to be involved into commercial trade. It most probably plays certain role as a natural controller of forestry pests, but due to low population density its influence on agriculture in this respect is negligible.	Terrestrial	No special conservation actions are known to date. Special research for revealing of possible cryptic taxonomic diversity and clarifying of the species natural habits are recommended.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Barbastella		darjelingensis	Hodgson [ in Horsfeld]	1855	0	Ann. Mag. Nat. Hist.	ser. 2, 16: 102	Darjeeling Barbastelle	None.		Taiwan; Sichuan China, N India, Nepal	Not listed.	Least Concern	Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).	Barbastella darjelingensis	1005652	23	Eastern Barbastelle	Asian Barbastelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PLECOTINI	Barbastella	NA	darjelingensis	Hodgson	1855	1	Plecotus_darjelingensis	Hodgson, B. H. (1855). In Horsfield, T. Brief notices of several new or little-known species of mammalia, lately discovered and collected in Nepal, by Brian Houghton Hodgson, Esq. Annals and Magazine of Natural History, Ser. 2, 16, 103.	https://www.biodiversitylibrary.org/page/2263778#page/130/mode/1up	BM 1854.9.1.13		Darjeeling, India.			darjelingensis (Hodgson, 1855)|dargelinensis Dobson, 1875|blanfordi Bianchi, 1917	split from B. leucomeles	Zhang, J. S., Han, N. J., Jones, G., Lin, L. K., Zhang, J. P., Zhu, G. J., ... & Zhang, S. Y. (2007). A new species of Barbastella (Chiroptera: Vespertilionidae) from north China. Journal of Mammalogy, 88(6), 1393-1403.				Afghanistan|Pakistan|India|Nepal|Bhutan|Bangladesh|Myanmar|Laos|Vietnam|China|Taiwan	Asia	Palearctic	LC	0	0	0	Barbastella_darjelingensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Barbastella_darjelingensis	1005652	23	Eastern Barbastelle	Asian Barbastelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Plecotini	Barbastella	NA	darjelingensis	B. H. Hodgson in Horsfield	1	Plecotus darjelingensis	Horsfield, T. 1855-08-01. Brief notices of several new or little-known species of mammalia, lately discovered and collected in Nepal, by Brian Houghton Hodgson, Esq. Annals and Magazine of Natural History (2)16(92):101-114.	https://www.biodiversitylibrary.org/page/18661014	BMNH:Mamm:1854.9.1.13	syntypes	https://data.nhm.ac.uk/object/44442bd3-afec-4c63-a2e5-27f764e33574	Darjeeling, India.	27.03333	88.26667	split from B. leucomeles	Zhang, J. S., Han, N. J., Jones, G., Lin, L. K., Zhang, J. P., Zhu, G. J., ... & Zhang, S. Y. (2007). A new species of Barbastella (Chiroptera: Vespertilionidae) from north China. Journal of Mammalogy, 88(6), 1393-1403.				Afghanistan|Pakistan|India|Nepal|Bhutan|Bangladesh|Myanmar|Laos|Vietnam|China|Taiwan	Asia	Palearctic	LC	0	0	0	Barbastella_darjelingensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Barbastella		darjelingensis	Hodgson [in Horsfeld]	1855	1	Ann. Mag. Nat. Hist.	ser. 2, 16: 102	Darjeeling Barbastelle	None.		Taiwan; Sichuan China, N India, Nepal	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85197261/85197270/' target='_blank'>Least Concern</a>	Distinct from leucomelas; see Zhang et al. (2007) and Benda et al. (2008). Does not include caspica, blanfordi,  or walteri; see Kruskop (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Barbastella darjelingensis; Barbastella darjelingensis; Barbastella darjelingensis; Barbastella darjelingensis; Barbastella darjelingensis; darjelingensis; dargelinensis; blanfordi; Barbastelle d'Asie; Ostliche Mopsfledermaus; Barbastelaasiética; Asian Barbastelle; Eastern Barbastelle; Asian Barbastelle; Darjeeling Barbastelle; B. darjelingensis